This is an archived copy of the 2020-2021 catalog. To access the most recent version of the catalog, please visit http://coursecatalog.bucknell.edu.

Biology (BIOL)

Biology is the natural science that concerns itself with study of the living world. The faculty of the biology department approaches the principles of the science from the unifying perspective of the theory of evolution. Emphases include the theory and practice of the way scientific investigations are conducted as well as the more practical applications of biology.

A major in Biology may serve as a sound preparation for those interested in careers in the life sciences including those who go on to graduate or medical school. Majoring in Biology also adds to students’ understanding of the issues concerned with health, the environment, and agriculture. In addition, Bucknell’s Biology majors are given the opportunity to become broadly educated “whole” scientists. They are encouraged to explore their interests within the humanities and social sciences.

Biology Majors

Two degree programs are offered through the biology program.

Bachelor of Arts

The Bachelor of Arts major requires eight courses in biology.

Core Sequence

BIOL 205Introduction to Molecules and Cells 11
BIOL 206Organismal Biology 11
BIOL 207Genetics 11
BIOL 208Principles of Ecology and Evolution 11
Four 300-level biology electives 24
CHEM 205Principles of Chemistry 31
or CHEM 207 Explorations in Chemistry
CHEM 211Organic Chemistry I 31
MATH 201Calculus I1
or MATH 202 Calculus II
MATH 216Statistics I1

Area I - Cellular/Molecular

BIOL 302Microbiology1
BIOL 304Biology of Cancer1
BIOL 323Mammalian Histology1
BIOL 324Neurophysiology1
BIOL 326Cytogenetics1
BIOL 327Molecular Biology1
BIOL 331Genomics1
BIOL 332Developmental Neurobiology1
BIOL 340Biochemical Methods1
BIOL 347Virology1
BIOL 348Immunology1
BIOL 352Cell Biology1
BIOL 363Receptors of Biological Membranes1
BIOL 365Introduction to Microscopy1
BIOL 375Cellular and Molecular Neurobiology1

Area II - Organismal

BIOL 309Wildlife and Emerging Diseases1
BIOL 312Comparative Vertebrate Anatomy1
BIOL 313Mammalogy1
BIOL 314Amphibian Biology and Conservation1
BIOL 316Plant Growth and Development1
BIOL 318Principles of Physiology1
BIOL 328Endocrinology1
BIOL 337Biology of Aging1
BIOL 339Developmental Biology1
BIOL 342Neuroethology1
BIOL 357Ornithology1
BIOL 358Invertebrate Zoology1
BIOL 359General Entomology1
BIOL 373Mycology1

Area III - Ecological/Evolutionary

BIOL 307Conservation Genetics1
BIOL 321Behavioral Ecology1
BIOL 330Plant Systematics1
BIOL 334Limnology1
BIOL 341Evolution1
BIOL 351Field Botany1
BIOL 353Ecosystem Ecology1
BIOL 354Tropical Ecology1
BIOL 355Social Insects1
BIOL 360Applications of Calculus to Medicine and Biology1
BIOL 361Systematic Biology1
BIOL 370Primate Behavior and Ecology1
BIOL 371Field Entomology1

Bachelor of Science

The Bachelor of Science major requires nine courses in Biology. The major provisions in Biology are the same as those noted under the Bachelor of Arts major, but five rather than four 300-level electives are required, only one of which may be BIOL 399 Mentored Undergraduate Research.

The Bachelor of Science major also requires:

CHEM 205Principles of Chemistry 41
or CHEM 207 Explorations in Chemistry
CHEM 211Organic Chemistry I 41
MATH 201Calculus I1
or MATH 202 Calculus II
MATH 216Statistics I1
PHYS 211
PHYS 212
Classical and Modern Physics I
and Classical and Modern Physics II
2
Select two of the following: 5
Biomimetic Materials
Organic Chemistry II (highly recommended)
Principles of Chemistry 2
Analytical Chemistry
Synthetic Organic Chemistry
Mechanistic Organic Chemistry
Special Topics in Organic Chemistry
Inorganic Chemistry I
Biochemistry I
Biochemistry II
Introduction to Computer Science
Data Structures & Algorithms
Environmental Pollution and Control
Environmental Planning
Introduction to Sustainable Design
Sustainable Resource Management
Applied G.I.S.
Food and the Environment
Global Change - Past and Present
Physical/Environmental Geology
Evolution of the Earth
Environmental GIS
Introduction to Geochemistry
Geomorphology
Paleontology
Psycholinguistics
Language and the Brain
Calculus II
Calculus III
Statistics II
Logic
Symbolic Logic
Philosophy of Science
Philosophy of Biology
Learning
Human Cognition
Biopsychology
Sensation and Perception
Comparative Animal Cognition
Advanced Psychological Statistics

Students interested in behavioral aspects of biology may wish to consider the Animal Behavior major; those interested in biochemistry, the Cell Biology/Biochemistry major; those interested in environmental issues, the Environmental Science bachelor of arts within the Environmental Studies program; and those interested in neurobiology, the Neuroscience program. Students planning to continue with graduate training in biology are encouraged to elect MATH 217 Statistics II and/or MATH 202 Calculus II, and to consult their academic adviser or pre-health professions adviser.

The recommended sequence for the Bachelor of Science major is as follows:

First Year
First SemesterCreditsSecond SemesterCredits
BIOL 2051BIOL 2061
CHEM 205 or 2071CHEM 2111
MATH 20161MATH 2161
Foundation Seminar1Elective1
 4 4
Sophomore
First SemesterCreditsSecond SemesterCredits
BIOL 2071BIOL 2081
Related area course71Related area course1
Elective1Elective1
Elective1Elective1
 4 4
Junior
First SemesterCreditsSecond SemesterCredits
Elective in biology1Elective in biology1
PHYS 2111PHYS 2121
Elective1Elective1
Elective1Elective1
 4 4
Senior
First SemesterCreditsSecond SemesterCredits
Two electives in biology2Elective in biology1
Elective1Elective1
Elective1Elective1
 Elective1
 4 4
Total Credits: 32

College Core Curriculum – Disciplinary Depth Requirements

Students in the biology major will satisfy the writing and the information literacy requirement by completing BIOL 205 Introduction to Molecules and Cells and BIOL 206 Organismal Biology and at least two 300-level biology laboratory/field courses. They will satisfy the formal presentation requirement by completing BIOL 206 Organismal Biology as well as at least two 300-level courses in biology, which will include a required oral presentation. The Culminating Experience in Biology requirement will be fulfilled by taking one 300-level laboratory or field course from a list provided by the department in one of a student's last three semesters.

Transfer students must complete at least four courses in biology in residence at Bucknell, only one of which may be BIOL 399 Mentored Undergraduate Research.

For Bucknell students who elect to study abroad, at least three upper division courses toward the major and at least one toward the minor must be taught by Bucknell faculty.

Students who pass the BIOL 121 Biology for Non-majors-BIOL 122 Biology for Non-majors sequence with a grade of B or better in both courses may receive one core credit toward the biology major pending consultation with the department chair.

Biology Minor

A minor in biology consists of five courses.

At least two 300-level courses 1
Select two of the following:
Introduction to Molecules and Cells
Organismal Biology
Genetics
Principles of Ecology and Evolution

Biology Department Learning Goals

(Numbers in parentheses are university-learning goals.)

I. Knowledge

Students completing the BA or BS major in Biology will:

  1. Demonstrate a proficiency with the concepts and methodologies of the traditional core of biology (1, 4, 6, 8, 9)
  2. Demonstrate the ability to use the scientific method (1, 8)
  3. Demonstrate the ability to apply concepts from diverse sub-disciplines of biology (1, 6, 8)
  4. Develop critical thinking skills to formulate and test biological questions (1, 4, 6)

In addition students completing a BS in biology will:

  1. Apply and integrate other related disciplines with their knowledge of Biology (1, 2, 6)

II. Communication

Students completing the BA or BS major in Biology will:

  1. Be able to write a well organized, logical and scientifically sound research paper or report (1, 7, 8)
  2. Be able to present a well organized, logical and scientifically sound oral presentation on a topic in Biology (1, 7, 8)

III. Career

Students completing the BA or BS major in Biology will:

  1. Be encouraged to pursue research experiences both on and off campus (1, 9)
  2. Be educated about the wide variety of career paths in Biology via seminars and invited speakers

Faculty in the department will provide strong personalized advising allowing students to make informed post-graduation decisions.

Non-STEM students fulfilling their natural sciences requirements will:

  1. Demonstrate a familiarity with biological principles and practices (4, 6, 8, 9)
  2. Appreciate the impact of biology on society and gain confidence in their ability to understand biologically related issues (4, 6, 8, 9)

Numbers in parentheses reflect related Educational Goals of Bucknell University.

Courses

BIOL 111. Controversies in Biology. 1 Credit.

Offered Occasionally; Lecture hours:3,Recitation:1
Introduction for the non-science major. Background on molecules, cells, and genetics. Required recitation will include discussions about current advances and controversies in biology. Not for pre-health students. Will not count toward the biology major. Students who take BIOL 111 may not take BIOL 121.

BIOL 120. Fight or Flight: The Biology of Stress. 1 Credit.

Offered Summer Session Only; Lecture hours:6
An exploration of biology through the lens of stress, this course will cover topics such as how stress relates to heart attacks, dwarfism, sex drive, memory loss, appetite, and aging. The course concludes with a biological-based discussion on how to effectively manage stress. Open to BCCSP.

BIOL 121. Biology for Non-majors. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Lab:3; May require dissection or live animal experimentation
Introductory course primarily for the non-biology major. Focuses on life at the cellular and biochemical levels, genetics, and biotechnology. This course is not appropriate preparation for the majority of pre-health graduate programs. Please consult with the Pre-health Adviser for more information.

BIOL 122. Biology for Non-majors. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Lab:3; May require dissection or live animal experimentation
Introductory course primarily for the non-biology major. Topics covered include principles of ecology, evolution, animal diversity, behavior, and structure, and function. This course is not appropriate preparation for the majority of pre-health graduate programs. Please consult with the Pre-health Adviser for more information.

BIOL 130. Health and Disease. 1 Credit.

Offered Occasionally; Lecture hours:3
A biology course, for non-science majors, that explores the basic biological principles underlying normal health and the most common diseases of humans. Students who have taken any 200-level Biology courses are not eligible for enrollment.

BIOL 131. Biology of Food. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:3
A course for non-majors that investigates fundamental concepts in biology through a focus on food and agriculture. We will debate current issues, such as genetic engineering, fad diets and our national farm policy. Lab involves hands on learning including growing and preparing food.

BIOL 150. Plants, People, and the Environment. 1 Credit.

Offered Fall Semester Only; Lecture hours:3
The diversity and evolution of plants, fungi, and related organisms with special emphasis on flowering plants; their importance for food, fiber, medicine, and psychoactive compounds; origins of agriculture; domestication of plants; and the role of plants in the environment.

BIOL 154. Society and the Environment. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
A biology course for non-majors only that explores society's impact on the environment and the environment's biotic and abiotic responses to various insults.

BIOL 1NT. Biology Non-traditional Study. .5-2 Credits.

Offered Fall, Spring, Summer; Lecture hours:Varies
Non-traditional study in Biology. Prerequisite: permission of the instructor.

BIOL 205. Introduction to Molecules and Cells. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:4
An introductory course which focuses on the molecular biology of cells. Basic biochemical processes, cellular and subcellular structure and function are emphasized. First core course.

BIOL 206. Organismal Biology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:4; May require dissection or live animal experimentation
An introductory course for biology majors emphasizing organisms as dynamic systems by integrating structure with function. Laboratories introduce scientific method and collaborative learning. Second core course.

BIOL 207. Genetics. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:1
A comprehensive survey of genetic mechanisms and methodologies, including classical genetics, recombinational analysis in bacteria, fungi, and higher eukaryotes, molecular genetics and populational and quantitative genetics. Third core course. Prerequisite: BIOL 205.

BIOL 208. Principles of Ecology and Evolution. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Lab:3
Introduction to systematic biology, evolutionary theory, physiological ecology, behavioral ecology, population and community ecology, and ecosystem structure and function. Fourth core course. BIOL 206 and BIOL 207 strongly recommended as prerequisites.

BIOL 220. Human Anatomy & Physiology I. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Lab:2; May require dissection or live animal experimentation
Introduction to human anatomy and physiology. Emphasis on the relationship between structure/function of the integumentary, musculoskeletal, nervous, and endocrine systems. Overview of anatomical terminology, cellular and tissue structures, and chemistry will be discussed. Does not count toward the biology major. Lab involves cat dissection. Prerequisite: permission of the instructor.

BIOL 221. Human Physiology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Lab:2
A course that focuses on the functions of and interactions between human organ systems. Does not count toward the biology major. Prerequisite: permission of the instructor.

BIOL 235. Microbiology for Health Professions. 1 Credit.

Offered Fall, Spring or Summer; Lecture hours:3,Other:3
This course will introduce students to the diversity of microorganisms by covering properties of eukaryotic and prokaryotic organisms, microbial genetics and biochemistry, and roles of microbes in disease. Labs will provide experience with common microbiology lab techniques: microscopy, identification, sterile technique, cell culture and plating, and staining methods.

BIOL 245. Tropical Marine Biology. 1 Credit.

Offered Summer Session Only; Lecture hours:10,Other:18
A field course in marine biology of coral reefs in the Virgin Islands for non-science majors. Prerequisite: permission of the instructor.

BIOL 254. Introduction to Ecology in West Africa. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:6
Introductory-level course in tropical ecology in West Africa, focusing on ecological phenomena with practical implications in Ghana. The course will examine basic ecological principles (explanations of biodiversity patterns, biological productivity, plant-animal interactions, habitat structures) and tropical habitats (rain forest, savannah, coastal lagoon, agricultural systems). Designed for non-science majors.

BIOL 266. Animal Behavior. 1 Credit.

Offered Both Fall and Spring; Lecture hours:3
A survey of important theories, issues, and empirical techniques in the interdisciplinary field of animal behavior emphasizing both proximate and ultimate mechanisms and explanations for behavior. Crosslisted as ANBE 266 and PSYC 266.

BIOL 2NT. Biology Non-traditional Study. 1-2 Credits.

Lecture hours:Varies,Other:Varies
Nontraditional study in biology.

BIOL 302. Microbiology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Lab:4
Ultra-structure, behavior, metabolism, molecular biology, and development of micro-organisms. Roles in disease and food production. Laboratory will emphasize cultivation and identification. Prerequisites: BIOL 205 and BIOL 207, and permission of the instructor. Crosslisted as BIOL 602.

BIOL 304. Biology of Cancer. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
The study of the molecular and cellular mechanisms that create cancer. Prerequisites: BIOL 205, BIOL 207, and permission of the instructor. Crosslisted as BIOL 604.

BIOL 306. Biology of Host-Microbe Interactions. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Other:2
Through study of the primary literature, this course will investigate the relationship between animals and their microbes, including evolution of host-microbe relationships, the impact of microbes on human health, the techniques used to study the microbiota, the biology of the microbiome, and other recent advances in the field. Crosslisted as BIOL 606.

BIOL 307. Conservation Genetics. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Other:3
As biodiversity has quickly eroded for the past few centuries, some scientists argue that humans are causing the 6th mass extinction event. This course emphasizes the application of population genetics, molecular phylogenetics, and reproductive genetics to answering biological questions in wildlife conservation. Crosslisted as ANBE 307 and ANBE 607 and BIOL 607.

BIOL 308. Microbial Genetics. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
Course focuses on molecular genetics of bacteria and archaea and the use of genetic tools to answer questions in microbiology. Primary literature will be used extensively. Prerequisites: permission of the instructor, BIOL 205 and BIOL 207 are required; BIOL 208 or BIOL 327 strongly recommended. Crosslisted as BIOL 608.

BIOL 309. Wildlife and Emerging Diseases. 1 Credit.

Offered Alternating Fall Semester; Lecture hours:3
Biology of wildlife diseases, especially zoonoses (infections that jump to humans). Course will integrate popular and scientific sources. Prerequisites: BIOL 206 and permission of the instructor. Crosslisted as BIOL 609.

BIOL 312. Comparative Vertebrate Anatomy. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:3; May require dissection or live animal experimentation
Gross morphology with emphasis on functional and evolutionary modifications of animal structure. Gross dissection and techniques used in morphology. Prerequisites: BIOL 122 or BIOL 206 and permission of the instructor. Crosslisted as BIOL 612.

BIOL 313. Mammalogy. 1 Credit.

Offered Alternating Fall Semester; Lecture hours:3,Other:3; May require dissection or live animal experimentation
Biology of mammals, including evolution, classification, biodiversity, behavior, anatomy, physiology, ecology, and conservation. Lab will include specimen identification, preparation, and field studies. Prerequisite: permission of the instructor. Crosslisted as BIOL 613.

BIOL 314. Amphibian Biology and Conservation. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:3
The biology of amphibians, including classification, physiology, reproduction, ecology, evolution, and conservation. Laboratory section will include identification of amphibians and field work to identify conservation issues surrounding local amphibian populations. Prerequisites: BIOL 206, BIOL 208 or 208E and permission of the instructor. Crosslisted as ANBE 314 and BIOL 614 and ANBE 614.

BIOL 316. Plant Growth and Development. 1 Credit.

Offered Alternating Fall Semester; Lecture hours:3,Other:3
The physiological and molecular bases of growth and development at the organ, tissue, and cellular levels. Effects of environmental stimuli and hormones on gene expression and the resultant changes at higher levels of organization. Prerequisite: BIOL 205, BIOL 206, and permission of the instructor. Crosslisted as BIOL 616.

BIOL 318. Principles of Physiology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Lab:3
Emphasizes the breadth of physiology and explores physiological principles of animals from a cellular, organismal, medical, and ecological framework. Laboratory focuses on experimental design and independent research. Prerequisites: BIOL 205, BIOL 206 and permission of the instructor. Crosslisted as BIOL 618.

BIOL 319. Seminar. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Recitation:1; Repeatable
Topics vary. Crosslisted as BIOL 619.

BIOL 320. Seminar. 1 Credit.

Offered Either Fall or Spring; Lecture hours:Varies,Other:3; Repeatable
Topics vary. Crosslisted as BIOL 620.

BIOL 321. Behavioral Ecology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3
How have ecological selection pressures (generated by animals’ biotic and abiotic environments) shaped the fascinating diversity of animal behaviors? Topics include habitat choice, foraging behavior, defenses against predation, cooperation and competition, sexual selection, and parental care. Heavy emphasis on primary literature and experimental design. Prerequisites: BIOL 208 and instructor permission. Crosslisted as ANBE 321, ANBE 621 and BIOL 621.

BIOL 323. Mammalian Histology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Other:3
A detailed study of the microscopic architecture and associated physiology of mammalian cells, tissues and organ systems. Prerequisites: BIOL 205 and BIOL 206 and permission of the instructor. Crosslisted as BIOL 623.

BIOL 324. Neurophysiology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3; May require dissection or live animal experimentation
A course that will explore the different circuits and systems of the brain and the corresponding brain structures. The laboratory portion of the course will include dissection of brain specimens to better visualize the content presented in class. Prerequisites: BIOL 205 and permission of the instructor. NEUR 253 recommended. Crosslisted as BIOL 624.

BIOL 325. Evolutionary Genomics. 1 Credit.

Lecture hours:3,Other:3
An exploration of evolutionary questions using large sequencing databases, with an emphasis on developing strategies for computational sequence analysis. Includes review of the primary literature. No coding experience necessary. BIOL 207 required; BIOL 208 recommended, but not required.

BIOL 326. Cytogenetics. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:3
Study of chromosome structure, organization, aberrations, and behavior. Multiple eukaryotic systems will be considered with links to human disease. Perquisites: BIOL 205 and BIOL 207 and permission of the instructor. Crosslisted as BIOL 626.

BIOL 327. Molecular Biology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Lab:3
Synthesis of DNA, RNA, and protein, and the regulation of these processes in both prokaryotic and eukaryotic cells; laboratory experience in the manipulation and analysis of genes. Prerequisites: BIOL 205 and BIOL 207 and permission of the instructor. Crosslisted as BIOL 627.

BIOL 328. Endocrinology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Problem Session:2
Regulation and function of hormones and their receptors from molecular to organismal levels. Role of hormones in development, physiology, and behavior; endocrine disease. Prerequisites: BIOL 205 and permission of the instructor. Crosslisted as BIOL 628.

BIOL 330. Plant Systematics. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:4
Exploration of the diversity of plant life on Earth through lectures, labs, and field trips; includes biogeography, natural history, evolutionary relationships, ethnobotanical uses, and identification. Prerequisite: BIOL 206 or permission of the instructor. Crosslisted as BIOL 630.

BIOL 331. Genomics. 1 Credit.

Offered Occasionally; Lecture hours:3,Other:2
A computer research-based course in which students study the structure, content, expression and evolution of genomes. Prerequisites: BIOL 207 and permission of the instructor. Crosslisted as BIOL 631.

BIOL 332. Developmental Neurobiology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Recitation:1
Primary literature-based senior seminar on topics in developmental neurobiology. Prerequisites: BIOL 205, BIOL 207, and either BIOL 206 or NEUR 100, junior or senior status, and permission of the instructor. Crosslisted as BIOL 632 and NEUR 332.

BIOL 334. Limnology. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:3
The physical, chemical, and biological characteristics of fresh-water communities are studied. Prerequisites: BIOL 208 and permission of the instructor. Crosslisted as BIOL 634.

BIOL 337. Biology of Aging. 1 Credit.

Offered Fall Semester Only; Lecture hours:3
This course will explore questions in the biology of aging from a physiological, genetic, and evolutionary framework, with an emphasis on critical reading of primary literature. Prerequisites: BIOL 206 and permission of the instructor. Crosslisted as BIOL 637.

BIOL 339. Developmental Biology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:3; May require dissection or live animal experimentation
This course provides an introduction to early animal development with emphasis on the molecular, cellular and genetic mechanisms that drive the formation of the embryo. Prerequisites: BIOL 205 and BIOL 206 and permission of the instructor. Crosslisted as BIOL 639.

BIOL 340. Biochemical Methods. 1 Credit.

Offered Spring Semester Only; Lecture hours:2,Other:6
A course in laboratory techniques including cell fractionation and analysis of proteins and nucleic acids. Spectrophotometry, chromatography, centrifugation, electrophoresis, and methods of molecular cloning are emphasized. Prerequisites: BIOL 205 and CHEM 351 and permission of the instructor. Crosslisted as CHEM 358.

BIOL 341. Evolution. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
Survey of evolutionary processes, phenomena, and mechanisms. Topics covered may include natural selection, sexual selection, adaptation, evolutionary constraints, speciation, evolution and development, coevolution, behavioral evolution, and macroevolution. Prerequisites: BIOL 208 and permission of the instructor. Crosslisted as ANBE 341 and ANBE 641 and BIOL 641.

BIOL 342. Neuroethology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
A course that integrates neurobiology and behavior in natural contexts. Emphasis on signal detection, recognition, discrimination, localization, orientation, and the control of complex acts. Neuronal and hormonal mechanisms, ontogeny and evolution of behavior will be considered. Prerequisites: BIOL 206 or BIOL 208 and permission of the instructor. Crosslisted as ANBE 342 and ANBE 642 and BIOL 642.

BIOL 347. Virology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:2
The study of virus structure, genome organization, replication and host-interactions. Emphasis will be on animal and bacterial viruses. Prerequisites: BIOL 205, BIOL 207, and permission of the instructor. Crosslisted as BIOL 647.

BIOL 348. Immunology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:3; May require dissection or live animal experimentation
Development and function of the immune system in animals. The immune response in health and disease. Techniques in immunology. Prerequisites: BIOL 205 and BIOL 206 and permission of the instructor. Crosslisted as BIOL 648.

BIOL 350. Independent Study. 1 Credit.

Lecture hours:Varies,Other:3; Repeatable
Selected topics.

BIOL 351. Field Botany. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:1
Outdoor field experience in plant diversity and ecology. Excursions to natural areas focused on identification, community dynamics, and ecological interactions/adaptations. Prerequisites: BIOL 208 and permission of the instructor. Crosslisted as BIOL 651.

BIOL 352. Cell Biology. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Lab:3
Covers biomembranes, cell growth patterns, cell signaling, the cytoskeleton, cell organelles, and microscopic techniques. Laboratory includes experience with cell culture. Prerequisites: BIOL 205 and permission of the instructor. Crosslisted as BIOL 652.

BIOL 353. Ecosystem Ecology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Recitation:1
Interactions between organisms and the physical and chemical environment including nutrient cycling and energy flow, biogeochemistry, and temporal and spatial dynamics of ecosystems. Prerequisites: BIOL 208, junior or senior status, and permission of the instructor. Crosslisted as BIOL 653 and ENST 353.

BIOL 354. Tropical Ecology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
Introduction to tropical ecology including life history strategies of vertebrates and invertebrates, biodiversity management and conservation. Emphasis on class and individual projects, data collection, and journal keeping. Prerequisites: BIOL 208 and permission of the instructor. Crosslisted as ANBE 354 and ANBE 654 and BIOL 654.

BIOL 355. Social Insects. 1 Credit.

Offered Fall Semester Only; Lecture hours:3,Other:3
Evolution and genetics of social behavior, caste, communication in foraging and colony defense, queen and worker control over reproduction, social homeostasis and population dynamics. Occasionally may be taught as a laboratory science. Prerequisites: BIOL 208 and permission of the instructor. Crosslisted as ANBE 355. Juniors and seniors only.

BIOL 357. Ornithology. 1 Credit.

Offered Occasionally; Lecture hours:3,Other:3
The biology of birds, including evolution, behavior, anatomy, physiology, ecology, and conservation; lab trips focus on identification of birds in the field. Prerequisites: BIOL 206, BIOL 208 and permission of the instructor. Crosslisted as ANBE 357 and ANBE 657 and BIOL 657.

BIOL 358. Invertebrate Zoology. 1 Credit.

Offered Alternating Fall Semester; Lecture hours:3,Other:3
A survey of the animal phyla covering phylogenetic relationships, functional morphology, ecology, life histories, symbiosis, ontogeny, and behavior. Includes hands-on study of organisms in lab and field. Prerequisites: BIOL 206, BIOL 208, and permission of the instructor. Crosslisted as BIOL 658.

BIOL 359. General Entomology. 1 Credit.

Offered Alternating Fall Semester; Lecture hours:3,Other:3
The biology of insects and their kin: anatomy, physiology, ecology, behavior, development, evolution, systematics, and diversity. Prerequisites: BIOL 206, BIOL 208, and permission of the instructor. Crosslisted as BIOL 659.

BIOL 360. Applications of Calculus to Medicine and Biology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3
Biology has been described as the most mathematical science. Researchers in biology use mathematical models to design strategies for controlling epidemics, administering drugs, and managing ecosystems. In this class you will learn how to develop your own models, approximate solutions to your models, and compare these solutions to real data. Crosslisted as BIOL 662 or MATH 260.

BIOL 361. Systematic Biology. 1 Credit.

Offered Occasionally; Lecture hours:3
Seminar in systematics, the study of the classification, diversity, and evolutionary relationships of all life. Emphasis placed on molecular data and the importance of systematics to all fields of biology. Prerequisites: BIOL 207, BIOL 208 and permission of the instructor. Crosslisted as BIOL 661.

BIOL 362. Topics in Cell Biology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:1.5
Selected topics in cell biology will be covered in a format (lecture, discussion, demonstration, seminar) that may vary from year to year. Topics may include membrane structure and dynamics, cell signaling, the cytoskeleton, protein synthesis and targeting, the cycle cycle, mitosis, cell-cell interactions and cell-substrate interactions, among others.

BIOL 363. Receptors of Biological Membranes. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
A course focused on the receptors and channels that function in biological membranes. The primary research literature will be used to explore the molecular bases of cellular communication, neuronal connectivity, and sensory transduction. Prerequisites: BIOL 205 and either BIOL 206 or NEUR 253. Crosslisted as BIOL 663 and NEUR 363.

BIOL 364. Advanced Data Analysis in Biology. 1 Credit.

Offered Spring Semester Only; Lecture hours:2,Other:4
Data exploration and visualization using state-of-the-art computational techniques. Using “big data” from their own research projects or public transcriptomic datasets, students will learn to analyze/visualize complex biological datasets. Lab includes hands-on work with R/virtual reality. No prior programming experience required. Prerequisites: BIOL 205, BIOL 207, MATH 216, permission of instructor. Crosslisted as BIOL 664.

BIOL 365. Introduction to Microscopy. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:3
This course is designed as an overview of light and electron microscopy, with emphasis placed on the use of instrumentation. Prerequisite: Permission of the instructor. Crosslisted as BIOL 665.

BIOL 370. Primate Behavior and Ecology. 1 Credit.

Offered Fall Semester Only; Lecture hours:3; May require dissection or live animal experimentation
Introduction to research on prosimians, monkeys, and apes with emphasis on the evolutionary origin of diversity, habitat use, social structure, social behavior, and cognitive abilities. Prerequisites: BIOL 122 or BIOL 208 or ANBE 266 or BIOL 266 or PSYC 266 and permission of the instructor. Crosslisted as ANBE 370 and ANBE 670 and BIOL 670 and PSYC 370 and PSYC 670.

BIOL 371. Field Entomology. 1 Credit.

Offered Fall Semester Only; Lecture hours:2,Other:2
Introduction to insects in their natural habitats, with emphasis on insect collecting, taxonomy, identification, ecology, and natural history. Students will make a professional-quality insect collection and acquire skills appropriate for biodiversity surveys. Prerequisites: BIOL 208 and permission of the instructor. Crosslisted as BIOL 671.

BIOL 373. Mycology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Other:3
Biology of fungi, including evolution, classification, biodiversity, ecology, and medical implications. Lab will involve identification, microscopy, culturing techniques, and field collection. Prerequisite: BIOL 206. Crosslisted as BIOL 673.

BIOL 375. Cellular and Molecular Neurobiology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3,Lab:3
A course focused on the molecular and cellular mechanisms that drive the firing and wiring of neurons. Topics of emphasis include excitable membrane physiology, synaptic transmission, plasticity and learning, and sensory transduction. The required co-requisite laboratory component provides hands-on neurophysiology experience with live animal models. Crosslisted as BIOL 675.

BIOL 378. Evolutionary Medicine. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
This seminar course will explore evolutionary approaches to medical theory and practice, including topics such as the fundamental nature of and relationship between patients and disease, evolution of human defenses to illness, pathogen evolution, cardiovascular disease, cancer, reproductive medicine, and mismatches between the modern environment and the human body. Crosslisted as BIOL 678.

BIOL 379. Sensory Ecology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:4
Sensory Ecology will examine the evolutionary interplay of sensory systems, behavior, and the environment. Topics will include ultraviolet vision and foraging strategies, magnetic navigation, ultrasonic signaling, and electric communication. In lab, we will design and conduct field or lab-based experiments to test hypotheses of your choosing. Crosslisted as BIOL 679.

BIOL 399. Mentored Undergraduate Research. .5-2 Credits.

Offered Fall, Spring or Summer; Lecture hours:Varies,Other:Varies; Repeatable; May require dissection or live animal experimentation
Undergraduate research mentored by a faculty member. Prerequisite: permission of the instructor.

BIOL 3NT. Biology Nontraditional Study. .5-2 Credits.

Lecture hours:Varies,Other:Varies
Nontraditional study in biology.

Faculty

Professors: Elizabeth A. Capaldi, Kenneth A. Field (Chair), Mark F. Haussmann, Stephen D. Jordan, Christopher Martine (Associate Chair), Matthew E. McTammany, Leocadia V. Paliulis, DeeAnn M. Reeder, C. Tristan Stayton

Associate Professors: Z Morgan Benowitz-Fredericks, Julie A. Gates, Matthew B. Heintzelman, Marie C. Pizzorno, Mark D. Spiro, Emily Stowe, Mizuki Takahashi

Assistant Professors: Moria Cairns Chambers, Matthew Q. Clark, Jayne A. Kubat (Adjunct), Sarah Emily Lower

Visiting Assistant Professors: Cecilia Bove, James Pearson, Gwendolyn C. Williams

Laboratory Directors: Karin I. Knisely, Rebekah Stevenson, Kathryn B. Toner

Microscopy Specialist: Joseph G. Moore