Neuroscience (NEUR)

The program in neuroscience offers students an interdisciplinary major representing biology, psychology, animal behavior, chemistry, mathematics, biomedical engineering, and physics. The neuroscience major is intended to give students opportunities, through coursework and research experience, to study the nervous system, its development and influence on behavior (broadly defined). Our faculty are active and productive scholars who involve students in their research programs, and thus we view research experience as a key aspect to the learning process.

The neuroscience major is offered within the Bachelor of Science degree program. All students are strongly encouraged to participate in research with faculty, as volunteers in their laboratories, or through independent studies and honors theses. Faculty interests and facilities include cell and molecular wet labs, electroencephalography for studying brain activity and cognitive/affective and perceptual processes, animal behavior labs for studying behavior and development in vertebrates (we house four species of primates, as well as rats, fish, turtles, mice, prairie voles, and bats), and invertebrates (e.g., flies and honey bees). We also have facilities for studying vision, cognition, and hormones and behavior. Students who succeed in neuroscience will be well-equipped to go on to graduate study in neuroscience, biology, psychology, and medicine, as well as to work in a variety of other disciplines including fields relating to biotechnology, pharmaceuticals, or medical instrumentation.

Bachelor of Science In Neuroscience

The Bachelor of Science major in neuroscience requires 17 courses (12 basic courses, plus five required 300-level courses). The students choose these five courses from a specified list of advanced neuroscience courses. One of the five can be chosen from a list of related courses as described below.

Of the 17 total courses taken by neuroscience majors, the following 12 courses are required:

Program Requirements
NEUR 100Introduction to Neuroscience1
BIOL 205Introduction to Molecules and Cells1
BIOL 207Genetics1
NEUR/PSYC 250Biopsychology1
NEUR 253Neural Cell Biology1
CHEM 211
  & CHEM 212
Organic Chemistry I
   and Organic Chemistry II
MATH 201Calculus I1
MATH 216Statistics I1
Select one of the following:1
Developmental Psychobiology
Human Cognition
Sensation and Perception
Organismal Biology
PHYS 211
  & PHYS 212
Classical and Modern Physics
   and Classical and Modern Physics
Five advanced neuroscience electives 1,25

Advanced neuroscience electives must be chosen from the courses listed in Category I and Category II with a minimum of two from Category I and two from Category II. One credit of Independent Research, or one course from the neuroscience-related electives list can also be counted as one of the five. NEUR 399 Undergraduate Research is counted as lab credit course. Neuroscience-related research credit may be earned in ANBE, BIOL, CHEM, NEUR, or PSYC. See the Neuroscience Program Director for permission.


Students are encouraged to become involved in independent study research, such as NEUR 399 Undergraduate Research; however, only one undergraduate research credit can be counted toward the five additional courses required for the major. 

Category I

BIOL 320Seminar1
BIOL 324Neurophysiology1
BIOL/NEUR 332Developmental Neurobiology1
BIOL 342Neuroethology1
BMEG 441Neural Signals and Systems1
NEUR 348Behavioral Pharmacology1

Category II

PSYC 305Developmental Psychopathology1
PSYC 318Cognitive Aging1
PSYC 339Psychology of Music1
PSYC 340Behavioral Neuroscience1
PSYC 349Human Neuropsychology1
PSYC 352Face Perception1

Neuroscience-related Courses

General Neuroscience-related Courses
ANBE 391Research1
BIOL 318Principles of Physiology1
BIOL 328Endocrinology1
BIOL 337Biology of Aging1
BMEG 300Biotransport I1
BMEG 409Fabrication amd Experimental Design.5
BMEG 461Brain, Mind and Culture1
CHEM 375Undergraduate Research1
CHEM 376Undergraduate Research1
NEUR 344Developmental Brain Research1
NEUR 399Undergraduate Research1
PSYC 324Advanced Psychological Statistics1
PSYC 329Undergraduate Research1
Courses Reflect Interests In Cell and Molecular Neuroscience
BIOL 304Biology of Cancer1
BIOL 322Physiological Mechanisms1
BIOL 327Molecular Biology1
BIOL 331Functional Genomics1
BIOL 339Developmental Biology1
BIOL 340Biochemical Methods1
BIOL 347Virology1
BIOL 352Cell Biology1
CHEM 351Biochemistry I1
Courses Recommended but Not Required
BIOL 327Molecular Biology1
Select one of the following:
General Chemistry
   and General Chemistry
Inorganic Chemistry I
   and Analytical Chemistry
MATH 202Calculus II1

Students need not choose any specific course of study or concentration within neuroscience. However, students with interests in particular aspects of the neuroscience major may consider choosing among courses that reflect these interests, such as cognitive and behavioral neuroscience or cellular and molecular neuroscience. Students interested in pursuing such interests should consult with their adviser on how best to accomplish their goals.

The recommended sequence for the neuroscience Bachelor of Science major is as follows:

First Year
First SemesterCreditsSecond SemesterCredits
NEUR 1001PSYC 2501
BIOL 2051MATH 2161
MATH 2011 
 3 2
First SemesterCreditsSecond SemesterCredits
CHEM 2111CHEM 2121
BIOL 2071NEUR 248, PSYC 203, PSYC 204, PSYC 252, or BIOL 2061
NEUR 2531One 300-level advanced neuroscience elective 
 3 2
First SemesterCreditsSecond SemesterCredits
PHYS 2111PHYS 2121
One 300-level advanced neuroscience elective One 300-level advanced neuroscience elective 
 1 1
First SemesterCreditsSecond SemesterCredits
One 300-level advanced neuroscience elective One 300-level advanced neuroscience elective 
 0 0
Total Credits: 12

We recommend that students who come to Bucknell with Advanced Placement (AP) credit in calculus begin their CHEM sequence in the first semester of the first year.

Writing in the Major

Neuroscience students will satisfy the writing in the major requirement by completing BIOL 205 Introduction to Molecules and Cells. In BIOL 205 Introduction to Molecules and Cells students are given direct instruction by the professor and through assigned readings from a text on scientific writing. Students will submit sections of four laboratory reports describing the results and conclusions from their experiments. Students will receive feedback on preliminary drafts of each report and will be given an opportunity to discuss their drafts as they prepare their final submissions.

Formal Presentation Experience

Students in the NEUR major will satisfy the formal presentation requirement by completing NEUR 253 Neural Cell Biology. Other NEUR courses also offer instruction on, and assessment of, formal presentations, or leading discussions. Students who are interested in gaining further instruction on presentations might consider taking one or some of the following courses:

ANBE/BIOL 342Neuroethology1
BIOL 320Seminar1
BIOL 324Neurophysiology1
NEUR 248Developmental Psychobiology1
NEUR/PSYC 305Developmental Psychopathology1
NEUR/PSYC 348Behavioral Pharmacology1
PSYC 203Learning1
PSYC 315Language Development1
PSYC 318Cognitive Aging1
PSYC 339Psychology of Music1

Information Literacy

Neuroscience students will satisfy their information literacy requirement by completing BIOL 205 Introduction to Molecules and Cells and NEUR 253 Neural Cell Biology. Students may also satisfy their information literacy requirement by completing an independent study (NEUR 399 Undergraduate Research). Students who want their independent study credit to count toward the NEUR major should be prepared to document how their independent study addresses some aspect of neuroscience so as to warrant 300-level NEUR credit. In all these experiences students receive direct instruction on the gathering and assimilation of scientific literature through a variety of search mechanisms, including, for example, PubMed, Medline and PSYCinfo.

Culminating Experience

Neuroscience seniors will satisfy their Culminating Experience requirement through any one of the following activities:

  • Registering for Independent Research NEUR 399 Undergraduate Research in their senior year (for a minimum of .5 credit). Students who do so will be graded on their participation and competency throughout the semester and will also be required to submit a written laboratory report based on their practical experiences. As noted above, the content of the independent study should be directly related to some aspect of neuroscience.
  • Students will have the opportunity to register for NEUR 400 Senior Seminar in Neuroscience – a seminar series – (.25 credit) in the spring semester of their senior year. This seminar series will require attendance at no fewer than four lectures. For example: attending lectures given by invited speakers from Biology, Chemistry, Physics or Psychology would fulfill this requirement. Attendance will be mandatory and students will be required to submit a written summary/reaction of each lecture, which will be graded by the members of the NEUR faculty.


NEUR 100. Introduction to Neuroscience. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
A survey of the study of the nervous system and its structure and function, ranging from molecular analyses of neurons to electrical and other correlates of human cognition.

NEUR 248. Developmental Psychobiology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
Addresses development in humans from conception through adolescence with some comparative analysis with non-humans. Emphasis on both normal and atypical cognitive, neuropsychological and neurobiological development. Prerequisite: PSYC 100 or NEUR 100. Crosslisted as PSYC 248.

NEUR 250. Biopsychology. 1 Credit.

Offered Both Fall and Spring; Lecture hours:3
Biological bases of behavior and their relationship to motivation, learning, and perception. Prerequisite: one of the following: NEUR 100, PSYC 100, BIOL 206, ANBE 266 or permission of the instructor. Crosslisted as PSYC 250.

NEUR 253. Neural Cell Biology. 1 Credit.

Offered Fall Semester Only; Lecture hours:3
A core course for neuroscience focused on structure/function relationships in neural cells. Basic protein biochemistry, ion channel activity, protein receptors, cell signaling, electrical properties and response patterns will be emphasized. Recommended for sophomores. Prerequisites: BIOL 205 and PSYC 250 and permission of the instructor.

NEUR 305. Developmental Psychopathology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
Addresses the behavioral phenotypes of a variety of neurodevelopmental and neuropsychiatric disorders in the context of theories and processes of normal development. Genetic and neurobiological underpinnings of disorders are discussed. Prerequisites: NEUR 248 or PSYC 248 and permission of the instructor. Crosslisted as PSYC 305.

NEUR 313. Researching Behavioral Neuroscience. 1 Credit.

Offered Both Fall and Spring; Lecture hours:3
Following a general orientation to behavioral genetics and pharmacology, including basic techniques of mouse handling, we will conduct group experiments. In addition, each student will develop and conduct an independent research project. Prerequisites: PSYC 215 or MATH 216 and PSYC 250 or an applied research methods course and permission of the instructor. May be crosslisted as PSYC 313.

NEUR 319. Topics in Neuroscience. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3; Repeatable
Occasional seminars on selected topics of current interest in neuroscience. Prerequisites: BIOL 205, BIOL 207 and NEUR 100, junior or senior status and permission of the instructor.

NEUR 330. Neuroscience of Addiction. 1 Credit.

Offered Fall Semester Only; Lecture hours:3
A study of the changes in neurocircuitry and neurobiology that occur in the brain due to drugs of abuse and addiction. Prerequisites: NEUR 100 and BIOL 205 or permission of the instructor.

NEUR 332. Developmental Neurobiology. 1 Credit.

Offered Spring Semester Only; Lecture hours:3,Other:1
Primary literature-based senior seminar on topics in developmental neurobiology. Prerequisites: BIOL 205, BIOL 207, and either BIOL 206 or NEUR 100, junior or senior status, and permission of the instructor. Crosslisted as BIOL 332.

NEUR 340. Behavioral Neuroscience. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
Advanced study of the relationship between the brain and behavior. Seminar discussion of complex problems in the field of behavior neuroscience including genetics, mood disorders, drug abuse, cognition and consciousness. Crosslisted as PSYC 340. Prerequisite: PSYC 250 or permission of the instructor.

NEUR 344. Developmental Brain Research. 1 Credit.

Offered Spring Semester Only; Lecture hours:3; Repeatable
Students learn a variety of assessment techniques in developmental neuropsychology and neuroscience (including EEG) and conduct quantitative research culminating in written and oral reports. Crosslisted as PSYC 344. Prerequisite: permission of the instructor.

NEUR 348. Behavioral Pharmacology. 1 Credit.

Offered Either Fall or Spring; Lecture hours:3
Focus on drugs that affect the nervous system, drugs of abuse, therapeutic drugs, drug action, behavioral changes as a result of long-term drug use, animal models and human studies. Prerequisites: PSYC 250 or BIOL 205 and permission of the instructor. Crosslisted as PSYC 348.

NEUR 360. Honors Thesis. 1 Credit.

Offered Both Fall and Spring; Lecture hours:Varies,Other:15; Repeatable
Prerequisite: permission of the department.

NEUR 399. Undergraduate Research. .5-2 Credits.

Offered Either Fall or Spring; Lecture hours:Varies,Other:Varies; Repeatable
Research on any aspect of neuroscience. Research topics may be posed by students or faculty. Prerequisite: permission of the instructor.

NEUR 3NT. NEUR Non-traditional Study. 1-2 Credits.

Offered Fall, Spring, Summer; Lecture hours:Varies,Other:Varies
Non-traditional study course in neuroscience. Prerequisite: permission of the instructor.

NEUR 400. Senior Seminar in Neuroscience. .25 Credits.

Offered Spring Semester Only; Lecture hours:1
NEUR majors may elect to attend a lecture series in the spring semester to satisfy the Culminating Experience requirement. Students will prepare written reactions to each seminar, graded as pass/fail. Prerequisites: senior status and NEUR majors and permission of the instructor.


Director: Peter G. Judge

Affiliated Faculty: Mitchell I. Chernin, Elizabeth A. Capaldi Evans, Judith E. Grisel, Andrea R. Halpern, Peter G. Judge, James E. Lavine, Heidi Lorimor, Aaron Mitchel, Kevin P. Myers, Marie C. Pizzorno, Ruth Tincoff, Joseph V. Tranquillo, T. Joel Wade

Coordinating Committee: David W. Evans, Elizabeth C. Marin, Kathleen C. Page, Jennifer Rice Stevenson, Eric S. Tillman